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Early in the Second Age of Middle-earth, Elven smiths forge nine Rings of Power for mortal Men, seven for the Dwarf-Lords, and three for the Elf-Kings. Soon after, the Dark Lord Sauron makes the One Ring, and uses it to attempt to conquer Middle-earth. After defeating Sauron, Prince Isildur takes the Ring, but after he is killed by Orcs, the Ring lies at the bottom of the river Anduin for over 2,500 years. Over time, Sauron captures the Nine Rings and transforms their owners into the Ringwraiths. The One Ring is discovered by Déagol, whose kinsman, Sméagol, kills him and takes the Ring for himself. The Ring twists his body and mind, and he becomes the creature Gollum (Peter Woodthorpe) who takes it with him into the Misty Mountains. Hundreds of years later, Bilbo Baggins (Norman Bird) finds the Ring in Gollum's cave and brings it back with him to the Shire.

Decades later, during Bilbo's birthday celebration, the Wizard Gandalf (William Squire) tells him to leave the Ring for his nephew Frodo (Christopher Guard). Bilbo reluctantly agrees, and departs for Rivendell. Seventeen years pass, during which Gandalf learns that evil forces have discovered that the Ring is in the possession of a Baggins. Gandalf meets Frodo to explain the Ring's history and the danger it poses, and Frodo leaves his home, taking the Ring with him. He is accompanied by three Hobbits, his cousins, Pippin (Dominic Guard), Merry (Simon Chandler), and his gardener Sam (Michael Scholes). After a narrow escape from the Ringwraiths, the hobbits eventually come to Bree, from which Aragorn (John Hurt) leads them to Rivendell. Frodo is stabbed atop Weathertop mountain by the chief of the Ringwraiths, and becomes sickened as the journey progresses. The Ringwraiths catch up with them shortly after they meet the Elf Legolas (Anthony Daniels); and at a standoff at the ford of Rivendell, the Ringwraiths are swept away by the river.

At Rivendell, Frodo is healed by Elrond (André Morell). He meets Gandalf again, after the latter escapes the corrupt wizard Saruman (Fraser Kerr), who plans to ally with Sauron but also wants the Ring for himself. Frodo volunteers to go to Mordor, where the Ring can be destroyed. Thereafter Frodo sets off from Rivendell with eight companions: Gandalf; Aragorn; Boromir (Michael Graham Cox), son of the Steward of Gondor; Legolas; Gimli (David Buck) the Dwarf, along with Pippin, Merry, and Sam.

Their attempt to cross the Misty Mountains is foiled by heavy snow, and they are forced into Moria. There, they are attacked by Orcs, and Gandalf falls into an abyss while battling a Balrog. The remaining Fellowship continue through the Elf-haven Lothlórien, where they meet the Elf queen Galadriel (Annette Crosbie). Boromir tries to take the Ring from Frodo, and Frodo decides to continue his quest alone; but Sam insists on accompanying him. Boromir is killed by Orcs while trying to defend Merry and Pippin. Merry and Pippin are captured by the Orcs, who intend to take them to Isengard through the land of Rohan. The captured hobbits escape and flee into Fangorn Forest, where they meet Treebeard (John Westbrook). Aragorn, Gimli, and Legolas track Merry and Pippin into the forest, where they are reunited with Gandalf, who was reborn after destroying the Balrog.

The four then ride to Rohan's capital, Edoras, where Gandalf persuades King Théoden (Philip Stone) that his people are in danger. Aragorn, Gimli, and Legolas then travel to the Helm's Deep. Frodo and Sam discover Gollum stalking them in an attempt to reclaim the Ring, and capture him; but spare his life in return for guidance to Mount Doom. Gollum eventually begins plotting against them, and wonders if the giant spider Shelob might help. At Helm's Deep, Théoden's forces resist the Orcs sent by Saruman, until Gandalf arrives with the absent Riders of Rohan, destroying the Orc army.
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